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The formation of humic substances occurs through multiple pathways during the decay of plant and animal remains in soil. The main pathways are: (1) lignin modification, (2) polyphenols from lignin oxidation, (3) microbially produced polyphenols, and (4) sugar-amine condensation. While the lignin pathway was once favored, most investigators now believe polyphenols and quinones produced from lignin or microbially are the major precursors that polymerize with amino compounds to form humic substances. The speci
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