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Integrated signatures of secondary microbial gas within gas hydrate reservoirs: A case study in the Shenhu area, northern S ... 1 1 Introduction 1 2 Geologic setting 2 3 Samples and methods 2 3.1 Sample collection 2 3.2 Organic geochemical experiments 3 3.2.1 Gas geochemical analyses 3 3.2.2 TOC measurement 3 3.2.3 Bitumen extraction and biomarker analysis 3 3.3 Archaeal diversity analysis 4 4 Results 4 4.1 Gas composition and stable isotopic characteristics 4 4.2 Organic geochemical characteristics 4 4.2.1 TOC content 4 4.2.2 Biomarker distribution and characteristics 4 4.3 Archaeal diversity 5 5 Discussion 8 5.1 Origin of the hydrate decomposed gas 8 5.2 Biodegradation of thermogenic hydrocarbon gases within the gas hydrate reservoir 8 5.3 Biodegradation of the liquid thermogenic hydrocarbons within the gas hydrate reservoir 10 5.3.1 Origin of the liquid hydrocarbons within gas hydrate reservoir 10 5.3.2 Biodegradation of the allochthonous liquid thermogenic hydrocarbons 11 5.4 Implications from archaeal diversity within the in-situ sediments 11 5.5 Possible hydrocarbon sources for the gas hydrate accumulations 12 6 Conclusions 13 Declaration of competing interest 13 Acknowledgem

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