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Melvin Calvin and his colleagues discovered the Calvin cycle in the 1950s through experiments using radioactive carbon isotopes. They found that carbon dioxide is fixed into the three-carbon compound 3-phosphoglycerate within the chloroplasts of plants. The Calvin cycle then regenerates the five-carbon acceptor ribulose-1,6-bisphosphate through three stages: fixation, reduction, and regeneration. This cyclic process uses energy from ATP and NADPH produced in the light-dependent reactions to convert CO2 into

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